Crossword puzzle clue "that smarts"

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Here are the possible solutions for that smarts clue

Answer
Clue
Relevancy
aches
Clue
Smarts
Definition
ACHE, to endure a dull lasting pain, also AKE
acumen
Clue
Smarts
Definition
(Latin) mental keenness
astutely
Clue
With smarts
Definition
ASTUTE, shrewd and perceptive
brains
Clue
Smarts
Definition
BRAIN, to dash out the brains of
cardinalsbrains
Clue
Vatican College smarts?
dodo
Clue
Extinct bird not known for its smarts
Definition
(Portuguese) an extinct bird
footballiq
Clue
Dolphin's smarts
goodsense
Clue
Smarts

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