Crossword puzzle clue "they feature creatures"

They feature creatures

Creatures that can't move but have sex by constantly and simultaneously ejaculating on a full moon once a year. It's called "broadcast spawning" and there are videos and they are glorious

"Lord of the Rings" creatures that must speak in tiny, high-pitched voices when they're seeds

Ratlike feature, and a hint to a feature of the four starred answers

Creature feature feature

Pie feature, or feature of this puzzle's other four longest answers

Winged creatures from Ghana's capital?

Sci-fi creatures